HEAT oven to 350ºF.
COMBINE first 5 ingredients in large bowl. Mix cream cheese spread, lemon zest, juice and seasonings until blended. Add to noodle mixture; mix lightly. Spoon into 1-1/2-qt. casserole sprayed with cooking spray; cover.
BAKE 20 min. Meanwhile, mix cracker crumbs and butter.
STIR tuna mixture; top with crumb mixture. Bake, uncovered, 8 to 10 min. or until tuna mixture is heated through and topping is golden brown.